Deep-sea cephalopods    
Cephalopod Behavior & Ecology

Current Position

Doctoral Candidate, Department of Integrative Biology, University of California, Berkeley

Graduate Researcher/Collaborator, Midwater Ecology Group, Monterey Bay Aquarium Research Institute



University of California, Berkeley

Deep-sea Squid Defensive Behaviors

Advisor: Dr. Roy L. Caldwell

Expected date of graduation, May 2009



East/West Marine Biology Program, Northeastern University, Massachusetts

Intensive marine biology field- and course-work at 3 locations:

The Marine Science Center, Nahant, Massachusetts

Discovery Bay Marine Laboratory, Discovery Bay, Jamaica

Friday Harbor Laboratories, San Juan Island, Washington



Illinois Wesleyan University, Bloomington, Illinois

Bachelors of Science in Biology, Magna Cum Laude, May 2002



James Cook University, Townsville, Queensland, Australia

Study abroad concentrating on marine biology and statistics course-work


Course Designer and Lecturer - Natural History and Human Interactions in Monterey Bay, Osher Lifelong Learning Institute, California State University, Monterey Bay, 2009

Course Designer and Lecturer - The Deep Sea, Osher Lifelong Learning Institute, California State University, Monterey Bay, 2008

Graduate Student Instructor - Invertebrate Zoology, University of California, Berkeley, 2006

Graduate Student Instructor - Animal Biology: A Behavioral View, University of California, Berkeley, 2005

Graduate Student Instructor - General Biology, University of California, Berkeley, 2004, 2005

Teaching Assistant - Oceanography, Three Seas Program, Northeastern University, 2004

Teaching Assistant - Marine Conservation Biology, Three Seas Program, Northeastern University, 2004

Teaching Assistant and Lab Instructor - Marine Biology, Harvard University, 2003



Bush, SL, Robison, BH, & RL Caldwell. 2009. Behaving in the dark: locomotor, chromatic, postural and bioluimescent behaviors of the deep-sea squid Octopoteuthis deletron Young 1972. Biological Bulletin. 216: 7-22.

Bush, SL & BH Robison. 2007. Ink utilization by mesopelagic squid. Marine Biology. 152(3):485-494.

LaJeunesse, TC, Lee, S, Bush, SL, & JF Bruno. 2004. Persistence of non-Caribbean algal symbionts in Indo-Pacific mushroom corals released to Jamaica 35 years ago . Coral Reefs. 24(1):157-159.

Bush, SL, Precht, WL, Woodley, JD, & JF Bruno. 2004. Indo-Pacific mushroom corals found on Jamaican reefs.  Coral Reefs. 23(2): 234.

Fellowships, Scholarships, and Honors

National Science Foundation, Graduate Research Fellowship, 2006-2009
Outstanding Graduate Student Instructor, UC Berkeley, 2007

UC Museum of Paleontology Remington-Kellogg Grant, 2008
Robert and Nancy Beim Field Research Fellowship, 2008

Society of Integrative and Comparative Biology, Student Support, 2007
Graduate Research Funds, UC Berkeley, 2007, 2008

Sigma Xi Grants-in-Aid of Research, 2006
Julius Thomas Hansen Travel Fellowship, UC Berkeley, 2006

Graduate Student Summer Fellowship, UC Berkeley,  2006, 2007
Ralph I. Smith Grant for Marine Research, UC Berkeley, 2005

Bermuda Biological Station for Research, Course Scholarship, 2005
Phi Beta Kappa, National Honor Society, inducted 2002

Beta Beta Beta, Biology Honors Society, inducted 2002
Dean’s List, Illinois Wesleyan University, 1998-2000, 2001-2002

Golden Key International Honor Society, nominated 2001
Presidential Scholarship, Illinois Wesleyan University, 1998-2002


2004-present Researcher, deep-sea squid defensive behaviors. Ink release, ink chemical composition, marine organism reaction to cephalopod ink, deep-sea squid body patterning, deep-sea squid arm autotomy. Collaborator: Dr. Bruce Robison, Monterey Bay Aquarium Research Institute.

2007-present Researcher, liquid chromatography-mass spectroscopy of cephalopod ink. Collaborator: Dr. Ulla Anderon, University of California, Berkeley.

2006 Researcher, animal camouflage. Sulawesi, Papua, and Ambon, Indonesia.

2006 Assistant intern coordinator and researcher, deep-sea squid arm autotomy. Advisor: Dr. George Matsumoto, Monterey Bay Aquarium Research Institute.

2005 Researcher, stomatopod reaction to squid ink. Hawaii Institute of Marine Biology, Coconut Island.

2003 Intern, deep-sea squid ink release. Advisor: Dr. Bruce Robison, Monterey Bay Aquarium Research Institute.

2003 Researcher, courtship behavior in the Caribbean Reef Squid. Bonaire, Netherlands Antilles. Collaborator: Jennifer Mather, University of Lethbridge.

2003 Researcher, heat shock response in anemones. Independent study, Three Seas Program. Advisor: Dr. Michael Lesser, University of New Hampshire.

2003 Researcher, heterotrophy in corals. Discovery Bay, Jamaica. Independent study, Three Seas Program. Advisor: Dr. Pete Edmunds, California State University, Northridge.

2002 Researcher, primary productivity. Friday Harbor Laboratories, WA. Independent study, Three Seas Program. Advisor: Dr. Jan Newton, University of Washington.

2002 Researcher, bryozoan natural products. Advisor: Dr. Elizabeth Balser, Illinois Wesleyan University.

2001 Researcher, coral colonization. Orpheus Island Research Station, James Cook University, Australia. Advisor: Dr. Bette Willis, James Cook University.

2001 Station assistant. Orpheus Island Research Station, James Cook University, Australia.

Research Cruise Participant

R/V Nathaniel B. Palmer, Antarctica. MBARI, 2008, 2009
R/V Pt. Lobos, Monterey Bay. MBARI, 2003-present (64 cruises, including chief scientist)
R/V Western Flyer, Monterey Bay; Oregon. MBARI, 2003-present (12 cruises)
R/V Derek M. Bayliss, Monterey Bay. MBARI, 2006
R/V New Horizon, Gulf of California, Mexico. University of Rhode Island, 2006
R/V Pt. Sur, Monterey Bay. Moss Landing Marine Labs, 2008


Deep sea squid: What are they inking?
Science, Technology, Engineering, and Mathematics Colloquium, Hartnell College, 2009

How to survive as a squid in the deep sea
NOAA Ocean Explorer Teacher Enrichment Workshop, 2009

Hunting and hiding in the dark: Predator-prey interactions in the deep sea
Sea Creatures Rediscovered, Pacific Grove Natural History Museum, 2009

Deep-sea research: Understanding animal adaptations
California State University Channel Islands, Resource Management course at MBARI, 2008

Arm autotomy in a mesopelagic squid
Micheli Lab, Hopkins Marine Station, Stanford University, 2008

Deep-sea research and squid defenses
Lawrence Hall of Science MARE Summer Leadership Institute at MBARI, 2008

Animal adaptations to the deep sea
Camp SEA Lab at MBARI, 2007

Posing in the dark: Behavioral components of a deep-sea squid
World Congress of Malacology, Antwerp, Belgium, 2007

Defensive behavior of a deep-sea squid
American Society of Limnology & Oceanography, Santa Fe, New Mexico, 2007

Autotomy as a deep-sea squid defense
Society of Integrative & Comparative Biology, Phoeniz, Arizona, 2007

Why do deep-sea squid release ink in the dark?
American Malacological Society, Seattle, Washington, 2006

How does a midwater squid avoid becoming a sperm whale (or ROV) meal?
2006 MBARI Summer Intern Symposium

Ink release by deep-sea squid
11th International Deep-sea Biology Symposium, Southampton, England, 2006

It’s pitch-black down here: Why are the squids still inking?
2003 MBARI Summer Intern Symposium

Professional Service

Reviewer: Marine Biology Research

Reviewer: Journal of Experimental Marine Biology and Ecology

Reviewer: The Biological Bulletin

Educational, Outreach, and Volunteer Work

Kayaking & Snorkeling, Young Women in Science Program, Monterey Bay Aquarium

Squid Science: Field Notes from Stephanie Bush, UCMP web feature

Elkhorn Slough Foundation and National Estuarine Research Reserve, weeding of invasives

California Coastal Cleanup Day, Asilomar Beach

MBARI tour and seminar for Camp SEA Lab

Science Fair Judge, The Aptos Academy, 3rd grade

MBARI tour and seminar for the Lawrence Hall of Science MARE Summer Leadership Institute

MBARI tour and seminar for the Coastal Resource Management course, CSU Channel Islands

Professional Societies

American Academy of Underwater Sciences Scientific Diver
American Malacological Society
American Microscopical Society
American Society of Limnology and Oceanography
Animal Behavior Society
Divers Alert Network
Sigma Xi, Associate Member
Society for Integrative and Comparative Biology
Western Society of Naturalists
Women in Science UC Berkeley